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Breaking it down:
Cities follow special rules...
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We've seen how prepositions are used to describe motion to or away from places. But when the name of a city is involved, no preposition is used.
Cities were so important in the Roman world that they had their own special grammatical rules. The Romans thought of cities as unique, named locations that didn't need prepositions to show direction - the case ending alone was enough!
When expressing motion towards a city, use the accusative case with no preposition.
When expressing motion away from a city, use the ablative case with no preposition.
A further surprising rule is that when an action happens in a city, you do not use the preposition "in + ablative" as you would expect. Instead, a special case is used.
The locative is a special case that locates the action. In Latin, locus means "place".
For now, the only locative you need to recognise is Rōmae (in/at Rome).
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